en_tn_condensed/ezk/10/11.md

they went in any of their four directions

Possible meanings are 1) "their" refers to the creatures. AT: "they would go straight in any one of the four directions that the creatures looked towards" or 2) "their" refers to the wheels.

whatever direction the head faced, they followed

Possible meanings for "the head" are 1) this refers to the winged creatures. AT: "they went in whatever direction the winged creatures were looking" or 2) this refers to the front wheel. AT: "they followed in whatever direction the front wheel went"